PNGTrees – Levieria squarrosa Perkins
Barry Conn (NSW) & Kipiro Damas (LAE).
Copyright held by the authors, Royal Botanic Gardens and Domain Trust, and Papua New Guinea National Herbarium
Botanische Jahrbucher für Systematik, Pflanzengeschichte und Pflanzengeographie Vol. 52: 196 (1915)
Other Literature: W .Philipson, Flora Malesiana Series 1 Vol. 10:
Field Characters: Small trees 10-20 m high. Bole cylindrical 25 cm diam.; almost straight bole 15 m long; buttresses absent; spines absent; aerial roots absent; stilt roots absent. Bark grey or brown, almost smooth, slightly pustular; lenticels irregular; subrhytidome yellow or green; bark thickness <25 mm thick, 5.0 mm thick; blaze consisting of 2 layers; strongly aromatic; pleasant; outer blaze brown or yellow, with stripes, granular without splinters; inner blaze red or brown, with stripes, fibrous; exudate present, colourless, spotty, changing colour on exposure to air, to brown or orange, not sticky. Terminal buds not enclosed by leaves; complex hairs absent; stinging hairs absent; mature twig without hairs.
Leaves: Spaced along branches <internodes readily visible>. Leaves spiral, simple; petiole present 5-10 mm long, not winged, attached to base of leaf blade <not peltate>, not swollen; lamina broadest at or near middle, 4.0-9.0 (-11.5) cm long, 1.2-3.3 (-4.8) cm wide; lamina symmetric, margin shortly crenate usually indistinct, with 1 or 2, less often up to 10 small blunt teeth present or entire, not dissected or lobed, apex acuminate or obtuse, venation pinnate <secondary veins arising from the midrib along its length>, secondary veins open <spaced far apart to easily see tertiary veins>, not prominent, but visible, intramarginal veins absent; lower surface pale green or green, upper surface dark green subglossy, hairs absent; oil dots absent; domatia absent; stipules absent.
Flowers: Inflorescence axillary <from between a leaf and branch> and terminal <at branch ends>. Flowers arising from a single point and flowers on a branched axis flower clusters on long stalks (peduncles); flowers bisexual, not stalked, with many planes of symmetry <actinomorphic> perianth present, pale yellow; ovary superior <seated above petals and sepals; hypogenous>, carpels separate (when more than one) <apocarpous>, locules 16-17.
Fruits: Fruits arising from single point surrounded by a yellow to yellow-brown receptacle or fruits arranged on branched axis fruiting clusters on long stalks. Fruit 8-10 mm long, 5.0 mm diam., fruit green or orange, not spiny, fleshy, simple of separate carpels of the one flower, indehiscent, drupe.
Distribution: Eastern Highlands, East Sepik, Morobe, Western Highlands, Southern Highlands, Central & West Sepik.
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